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What two steps in oxidative phosphorylation can vary

Peter Mitchel eventually was awarded a Nobel Prize for understanding how oxidative
phosphorylation works. Most scientists didn’t accept his proposed mechanism because
there was not a strict quantitative relationship between reducing equivalent (electron)
consumption and ATP production as there was for all other known biological conversion
reactions.
A) What two steps (or molecular processes) in oxidative phosphorylation can vary in
efficiency so that this strict relationship is not maintained? (4 pts)

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